Val Eliot, a land girl working on an English farm during the war, finds herself in charge of protecting a rescued border collie named Peter Woodhouse from the owner who mistreated him. When Val meets Mike, an American Air Force pilot stationed nearby, she realizes that the safest place for the little dog is alongside Mike at the base. With the love and attention of the pilots, Peter Woodhouse becomes Dog First Class, the mascot of the U.S. Air Force, boldly accompanying the officers on their missions--until a disaster jeopardizes the future of them all. It is then that Peter Woodhouse brings Ubi, a German corporal, into their orbit, sparking a friendship that comes at great risk, but carries the richest of rewards.